Meet Irakli Loladze., Bryan College of Health Sciences

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Irakli Loladze

Irakli Loladze, PhD

PH: 402-481-8896

Email Irakli

Previous Education:

  • Postdoctoral, Princeton University
  • PhD, Arizona State University
  • MA, Arizona State University
  • BA, Tbilisi State University


I am a Mathematical Biologist

Notable publications:

Zhu C, Kobayashi K, Loladze I, Zhu J, Jiang Q, Xu X, Liu G, Seneweera S, Ebi KL, Drewnowski A, Fukagawa NK, Ziska, LH (2018) Carbon dioxide (CO2) levels this century will alter the protein, micronutrients, and vitamin content of rice grains with potential health consequences for the poorest rice-dependent countries. Science advances.  May 1; 4(5):eaaq1012.

(covered by Scientific American, National Post, Washington Post, and other media outlets)

The Impacts of Climate Change on Human Health in the United States: A Scientific Assessment (2016) Ziska LH, Crimmins A, Auclair,AN, DeGrasse SL, Khan AS, de Leon  BP, Thurston JA, Garofalo JD, Gendel SM, Loladze I, Showler AT, Walls I. Ch. 7 ‘Food Safety, Nutrition, and Distribution'

Ziska, LH, Pettis, JS, Edwards, J, Hancock, JE, Tomecek, MB, Clark, A, Dukes, JS, Loladze, I and Polley, HW (2016) Rising atmospheric CO2 is reducing the protein concentration of a floral pollen source essential for North American bees. Proc. R. Soc. B, 283:20160414. The Royal Society.

Deng, Q., Hui, D., Luo, Y., Elser, J.J., Wang, Y-P., Loladze, I., Zhang, Q., Dennis, S. (2015) Down-regulation of tissue N:P ratio in terrestrial plants grown under elevated CO2 levels. Ecology,

Loladze, I. (2014) "Hidden shift of the ionome of plants exposed to elevated CO2 depletes minerals at the base of human nutrition" eLife, 3, e02245 (F1000 rating: Exceptional)

Peace, A., Zhao, Y,. Loladze, I., Elser, J., Kuang, Y. (2013) A stoichiometric producer-grazer model incorporating the effects of excess food-nutrient content on consumer dynamics. Mathematical Biosciences, 244(2), 1879-3134,

Elser, J.J., Loladze, I., Peace, A.L., Kuang, Y. (2012) Lotka re-loaded: Modeling trophic interactions under stoichiometric constraints.  Ecological Modeling, 245, 3–1,

Loladze, I (2019) Iterative chemostat: A modelling framework linking biosynthesis to nutrient cycling on ecological and evolutionary time scales. Mathematical Biosciences and Engineering, 16(2): 990-1004. doi: 10.3934/mbe.2019046


Loladze, I., Nolan, J. M., Ziska, L., & Knobbe, A. (2018). Rising atmospheric CO2 tends to lower carotenoid levels in plants: Can the global decline in dietary carotenoids affect human vision and memory? Journal of Alzheimer's Disease, 64 (3), 1042.

Loladze, I. and Elser J.J.  (2011) The origins of the Redfield nitrogen-to-phosphorus ratio are in a homoeostatic protein-to-rRNA ratio, Ecology Letters, 14, 244-250  (Editor’s Choice, Science)

Loladze, I., Kuang, Y., Elser, J.J. & Fagan, W.F. (2004) Competition and Stoichiometry: Coexistence of two predators on one prey. Theoretical Population Biology 65, 1-15

Loladze, I. (2002) Rising CO2 and human nutrition: toward globally imbalanced plant stoichiometry? Trends in Ecology and Evolution, 17, 457-461 (highlighted in Nature)

Loladze, I., Kuang, Y. Elser, J.J. (2000) Stoichiometry in producer-grazer systems: linking energy flow with element cycling, Bulletin of Mathematical Biology 62, 1137-1162 (in the top ten cited articles in the BMB since 1999)

Professional Service and Recognition:

The Teaching Council of the UNL Award  (2006, 2007, 2008)

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